Center and surround

Every point of an image is seen by many overlapping receptive fields. Each one reports how its center differs from its surround. Each preset below runs the same machinery on a different pattern of light.

Demonstrations

Light on the photoreceptors

The stimulus. Click or drag on any panel to choose the recorded cell.

darklight

Horizontal cells: the surround

A wide, blurred average of the photoreceptor signal around each point.

belowadapted levelabove

ON bipolar cells

Center minus surround. Graded voltage, no spikes. OFF bipolar cells are the mirror image.

hyperpolarizedrestdepolarized

Ganglion cells

Firing rate. A rate cannot fall below zero; the maintained rate is 22 Hz.

0 Hz22 Hz maintained100 Hz

One row, read across the population

Every cell along the dashed row, left to right. The amber line marks the recorded cell.

The recorded cell

This cell's response as the spot grows

Arrow keys or Page Up / Page Down step through the demonstrations; 1–4 answer a prediction.